Royal Netherlands Institute for Sea Research
Phone number
+31 (0)222 36 9488
Senior Scientist
  • Ecology and Evolution, Zoology
  • Marine mammal ecology, dolphins, cetaceans
  • Seabird ecology, behaviour, fouraging
  • Marine protected areas, fisheries, oil pollution
  • Animal tracking, colour ringing, GPS tagging
UvA Bird Tracking System.

Dr. Kees Camphuijsen

Senior Scientist

Birds and mammals as part of a large jigsaw puzzle

Marine ecologist Kees Camphuysen obtained his PhD for his long-term research into the herring gulls and lesser black-backed gulls in colonies on the island of Texel. ‘These birds’ lives are strongly related to commercial fishing activities on the North Sea. The development of the chicks is synchronised with commercial fishing efforts in the vicinity. For example, the chicks scarcely grow on a Saturday or a Sunday, and cannibalism is high if the fishing boats are in harbour during weekends. Alternative sources of food, of which there are many, are insufficient to stave off this weekly recurring starvation. For the herring gull, there is a strong relationship with food supplies in cities. Since we have covered up landfill areas, somewhere in the 1990s, herring gulls turned their attention to our villages, towns and tourist centres. Natural resources are insufficient to maintain the populations which increased so strongly in the past. Consequently, the number of herring gulls has been decreasing for many years.’

Nuances of the North Sea

‘Ultimately, this sort of puzzle is just a small part of a far bigger picture. Research into birds requires a proper understanding of their environment, but also of their relations with other species. In our current research in the North Sea, we are trying to map and quantify all of those complex connections. This concerns interactions between species – many birds and sea mammals are mutually dependent in their search for food – but also the characteristics of the areas they forage in. Not everything that floats on the North Sea is water. Salt concentrations, depth, currents, seabed fauna and visibility are just a few of the factors that determine why one location can be highly suitable for a species, while another is not.’

Using birds to take a look below the surface

‘It is highly tempting for a biologist to use transmitters and other high-tech equipment to study not just one species but even a limited number of individuals. However, you then risk losing sight of the bigger picture, notably the species interactions. By no longer viewing bird research independently of the environment and the interactions between different species, we understand more about the impact of human interventions, such as the construction of offshore wind parks or offshore islands. As research below the water surface is relatively difficult, we combine seagoing research of the clearly visible birds and seals, for example, with demographical studies in their colonies or roosts on land. By simultaneously measuring their “performance” on land (e.g. survival, number and growth of the young), we hope to learn more about the quality of their living environment at sea.’



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Research interests

Marine studies and research interests (1984-present) include at-sea work in the Atlantic Ocean, studying offshore distribution patterns, ecological interactions and interactions with fisheries of charismatic megafauna (whales, dolphins, turtles, seals, large predatory fish, and seabirds). Areas of research include the Greenland Sea, Norwegian Sea, North Sea, Atlantic Ocean, NW African shelf waters, SW African shelf waters, South Georgia, South America, and the Antarctic Peninsula.

Seabird colony studies on the foraging ecology, demography and migratory movements of Herring Gull Larus argentatus and Lesser Black-backed Gulls Larus fuscus. Colony work commenced in 2005, colour-ringing programmes and diet studies since 2006, and GPS tracking since 2008.

Studies of marine oil pollution since early 1970s through systematic beached bird surveys within The Netherlands and impact assessments of major oil spills throughout Europe.

Recent research interests in collaboration with the Netherlands Institute for the Study of Crime and Law Enforcement (NSCR) focus on Illegal, Unreported and Unregulated fisheries worldwide.



Positions & Working groups

Senior scientist at Royal Netherlands Institute of Sea Research (NIOZ) 

Senior scientist Programme Biodiversity Oil and Gas, Mauritania

Secretary of the World Seabird Union 

Standing member European Seabirds at Sea database 

Former chairman Dutch seabird group 

Fellowship Nederlands Studiecentrum Criminaliteit en Rechtshandhaving (NSCR)

Editorial Duties

Member editorial board ICES Journal of Marine Science 

Member editorial board Seabird 

Former member editorial board Lutra 

Former member editorial board and former Technical editor Ardea 

Former editor in chief Atlantic Seabirds 

Former editor in chief Sula 


Key publications

Donk S. van, C.J. Camphuysen, J. Shamoun-Baranes & J. van der Meer 2017. The most common diet results in low reproduction in a generalist seabird. Ecology and Evolution DOI: 10.1002/ece3.3018.

Shamoun-Baranes J., J.B. Burant, E.E. van Loon, W. Bouten & C.J. Camphuysen 2016. Short distance migrants travel as far as long distance migrants in lesser black-backed gulls Larus fuscus. J. Avian. Biol. 47: doi: 10.1111/jav.01299.

Camphuysen C.J., J. Shamoun-Baranes, E.E. van Loon & W. Bouten 2015. Sexually distinct foraging strategies in an omnivorous seabird. Mar. Biol. 162: 1417-1428.

Camphuysen C.J., J. Shamoun-Baranes, W. Bouten & S. Garthe 2012. Identifying ecologically important marine areas for seabirds using behavioural information in combination with distribution patterns. Biol. Conserv. 156: 22-29.

Camphuysen C.J. & A. Gronert 2012. Apparent survival and fecundity of sympatric Lesser Black-backed Gulls and Herring Gulls with contrasting population trends. Ardea 100: 113-122.

Please find my list of publications at the bottom of this webpage. You can download all my publications on ResearchGate. See also: PhD-thesis


Professional education



Camphuysen C.J. & P.A. Henderson 2017. North sea fish and their remains. Royal Netherlands Institute for Sea Research & Pisces Conservation Ltd, Texel, The Netherlands, 325pp. ISBN 9781904690658. A summary of the book is to download here.

Camphuysen C.J. & M.L. Siemensma 2011. Conservation plan for the Harbour Porpoise Phocoena phocoena in The Netherlands: towards a favourable conservation status. Royal Netherlands Institute for Sea Research, Texel, 183pp DOI: 10.13140/RG.2.1.2099.9765.

Camphuysen C.J. & Peet G. 2006. Walvissen en dolfijnen in de Noordzee. Fontaine Uitgevers, Kortenhoef, 160pp ISBN-10:90 5956 2216

Camphuysen C.J. & Peet G. 2006. Whales and dolphins of the North Sea. Fontaine Uitgevers, Kortenhoef, 160pp ISBN-13:978 98 5956 2219

Boyd I.J., Wanless S. and Camphuysen C.J. (eds) Top predators in Marine Ecosystems: monitoring change in upper trophic levels. Cambridge Univ. Press, Cambridge.

Bijlsma R.G., Hustings F. & Camphuysen C.J. 2001. Algemene en schaarse vogels van Nederland. Avifauna van Nederland, 2. KNNV Uitgeverij Utrecht en GMB Uitgeverij, Haarlem, 497pp. ISBN 90-74345-21-2

Bird tracking systems

Refereed contributions to books

Camphuysen C.J. & C. Smeenk 2016. Noordkaper - Bruinvis. In total 27 book chapters In: Broekhuizen S., K. Spoelstra, J.B.M. Thissen, K.J. Canters & J.C. Buys (eds) 2016. Atlas van de Nederlandse Zoogdieren: 302-374. Natuur van Nederland 12, Naturalis Biodiversity Center, Eis Kenniscentrum insecten en andere ongewervelden, Zoogdiervereniging en KNNV Uitgeverij, Zeist.

Camphuysen C.J. & M. Heubeck 2015. Beached bird surveys in the North Sea as an instrument to measure levels of chronic oil pollution. In: Carpenter A. (ed.) Oil Pollution in the North Sea 193-208. Handbook of Environmental Chemistry, Springer-Verlag, Berlin, Heidelberg, New York.

Camphuysen C.J. & B. Vollaard 2015. Oil pollution in the Dutch sector of the North Sea. In: Carpenter A. (ed.) Oil Pollution in the North Sea: 117-140. Handbook of Environmental Chemistry, Springer-Verlag, Berlin, Heidelberg, New York.

Zuur A.F. & C.J. Camphuysen 2012. Generalised additive models applied on northern gannets. In: Zuur A.F. (ed.) A beginner's guide to generalised additive models with R: 145-168. Highland Statistics, Newburgh.

Camphuysen C.J. & de Leeuw, J. 2011. The impact of hydrophobic and insoluble chemicals released from merchant shipping on European marine ecosystems and wildlife. In: Calewaert J.B. & N. McDonough (eds) Chemical Pollution in Europe's Seas: Programmes, Practices and Priorities for Research, Marine Board Position Paper 16: 51-63. Marine Board-ESF, Ostend, Belgium.

Camphuysen C.J., B. Scott & S. Wanless 2006. Distribution and foraging interactions of seabirds and marine mammals in the North Sea: multi-species foraging assemblages and habitat-specific feeding strategies. In: Boyd I.J., Wanless S. & Camphuysen C.J. (eds) Top predators in Marine Ecosystems: monitoring change in upper trophic levels: 82-97. Cambridge Univ. Press, Cambridge.

Camphuysen C.J. 2001. Seabirds and fisheries interaction. In: Steele J.H., Thorpe S.A. & Turekian K.K. (eds) Encyclopedia of Ocean Sciences: 2677-2686. Academic Press, London.

Bourne W.R.P. & Camphuysen C.J. 2000. Seabirds. Chapter 114, In: Sheppard C. (ed.) Seas at the Millenium, an environmental evaluation: 105-116. Elsevier, Amsterdam.

Camphuysen C.J. & Garthe S. 2000. Seabirds and commercial fisheries: population trends of piscivorous seabirds explained? In: Kaiser M.J. & Groot S.J. de (eds). Effects of fishing on non-target species and habitats: Biological, Conservation and Socio-Economic Issues: 163-184. Blackwell Science, Oxford.

Camphuysen C.J. 1997. Ivory Gull Pagophila eburnea. In: Hagemeijer J.M. & Blair M. (eds) The EBCC atlas of European breeding birds: their distribution and abundance: 348. T. & A.D. Poyser, London.

Camphuysen C.J. 1994. Ivory Gull Pagophila eburnea. In: Tucker G.M. & Heath M.F. (eds). Birds in Europe - their conservations status: 290-291. Birdlife Conservation Series No. 3, Birdlife International, Cambridge.

Camphuysen C.J. 1985. Zeetrektellingen. In: Hustings M.F.H., Kwak R.G.M., Opdam P.F.M. & Reijnen M.J.S.M. (eds). Vogelinventarisatie: 215-219. Pudoc, Wageningen.

Linked news

Monday 03 October 2022
Bird flu goes offshore | Sudden spread of 'highly pathogenic' virus among seabirds and even marine mammals
After years of outbreaks among both domesticated and wild birds on land, the bird flu virus H5N1 has moved to sea. In addition to the massive outbreak among sandwich terns, from Texel to northern France, thousands of seabirds have perished in the…
Monday 22 August 2022
EU-Horizon Europe funding for ACTNOW project
Human activities have created unprecedented, cumulative threats resulting in stunning losses of biodiversity in our oceans. This is leading to well‐documented declines in seafood resources, losses of iconic and culturally valuable habitats, and…
Wednesday 17 August 2022
Exhibition draws attention to plastic pollution
Arnold Gronert spent years beachcombing near Petten (Noord-Holland), or looking for washed up birds and marine mammals. He made an enormous contribution to the strandings dataset, kept for decades at NIOZ and at Naturalis. The spectacular amounts of…
Wednesday 08 December 2021
Stranding of puffins in the Netherlands continues
In early December 2021, an unusual stranding of dead and weakened Atlantic Puffins Fratercula arctica started on the Dutch coast. Specimens washed ashore on all Wadden Islands, but also scattered along the mainland coast and in Zeeland. Because this…
Thursday 11 November 2021
Oceans of Energy will build 1 MW offshore solar off the coast of Scheveningen
Scheveningen, The Netherlands; November 11, 2021 - Today Oceans of Energy and partners & observers announce the upscaling of the world's first offshore solar farm system 20 times and expand it to 1 MW (MegaWatt). In the coming years, the company…
Thursday 29 October 2020
Zeer zeldzame Gaffelmakreel in de NIOZ fuik in het Marsdiep
Op 8 oktober 2020 werd een bijzondere vis aangetroffen in het Marsdiep (Texel) in de visfuik die het NIOZ al sinds 1960 gebruikt om veranderingen in de visfauna in de Waddenzee te volgen. Het ging om een Gaffelmakreel Trachinotus ovatus van bijna 32…
Friday 26 June 2020
Blog on gull research | Fish and chicks
UvA/NIOZ researcher Roos Kentie tells us about her fieldwork and research. 'A few nests of unfortunate gulls got predated, but most eggs of the Herring and Lesser Black-backed Gulls in our research colony on the south tip of Texel have hatched. Gull…
Friday 19 June 2020
The North Sea’s forgotten angels
The presence of Angelsharks in the North Sea has faded from our memory. In a recent publication in Marine Biology, WWF nature conservationists and NIOZ researchers urge that we put top predators from earlier centuries back into the ‘baseline’ picture…
Friday 10 January 2020
Zilvermeeuw vergroot broedsucces met huisvuil
De zilvermeeuwen die broeden op de zuidpunt van Texel krijgen meer en grotere jongen wanneer ze voedselzoeken op vuilnisbelten en in steden. Maar het zoeken van antropogeen – niet natuurlijk – voedsel heeft een nadeel: het kost meer energie met als…
Wednesday 18 December 2019
Olievervuiling Noordzee naar aanvaardbaar minimum teruggebracht
In tijden van zorg over een veranderend klimaat, de plasticsoep in zee, stikstofuitstoot en andere milieuproblemen wordt soms vergeten dat beleid ook effectief kan zijn. In een vandaag gepubliceerd rapport van het Koninklijk Nederlands Instituut voor…

Linked blogs

Monday 19 June 2023
ACTNOW Expedition June 2023
The 10 day cruise to sample the Doggers bank is organised in the framework of the EU project ACTNOW*. The aim is to collect data on different elements of the food web simultaneously: from phytoplankton (the small “plants” in the sea) via grazing…
Tuesday 05 January 2021
NIOZ@SEA | Audioblogs door Threes Anna vanaf RV Pelagia op de Noordzee
Threes Anna maakte een audioreportage in 4 delen van onderzoek op de Noordzee met marien ecoloog Kees Camphuysen. Een gesprek vanaf het dak van de brug van de RV Pelagia, waar ze 10 dagen van zonsopkomst tot zonsondergang waarnemingen vastleggen over…
Friday 26 June 2020
Gulls | Fish and chicks
UvA/NIOZ researcher Roos Kentie tells us about her fieldwork and research. 'A few nests of unfortunate gulls got predated, but most eggs of the Herring and Lesser Black-backed Gulls in our research colony on the south tip of Texel have hatched. Gull…
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Monday 13 May 2019
NIOZ@SEA | North Sea Expeditions 2019
In 2019 three Pelagia cruises will be organised to study the carrying capacity of the North Sea. Each cruise will last 2 to 3 weeks and follows different routes to cover and describe the various parts of the ecosystem of the North Sea. On May 8th the…

NIOZ publications

  • 2023
    Brown, J.M.; Bouten, W.; Camphuysen, C.J.; Nolet, B.A.; Shamoun-Baranes, J. (2023). Energetic and behavioral consequences of migration: an empirical evaluation in the context of the full annual cycle. NPG Scientific Reports 13(1): 1210.
    Camphuysen, C.J.; van Donk, S.; Shamoun-Baranes, J.; Kentie, R. (2023). The Annual Cycle, Breeding Biology and Feeding Ecology of the Lesser Black-Backed Gull Larus fuscus. Ardea 112(1).
    Furness, R.W.; Gear, S.C.; Camphuysen, C.J.; Tyler, G.; de Silva, D.; Warren, C.J.; James, J.; Reid, S.M.; Banyard, A. (2023). Environmental samples test negative for avian influenza virus H5N1 four months after mass mortality at a seabird colony. Pathogens 12(4): 584.
    Kentie, R.; Morgan Brown, J. ; Camphuysen, C.J.; Shamoun-Baranes, J. (2023). Distance doesn't matter: migration strategy in a seabird has no effect on survival or reproduction. Proc. - Royal Soc., Biol. Sci. 290(1997): 20222408.
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    ten Horn, J.; Kühn, S.; Camphuysen, C.J.; de Groot, H.; Keijl, G.O. (2023). Roodsnavelkeerkringvogels Phaethon Aethereus in Nederland. Sula 31: 1-12
  • 2022
    Brown, J.M.; Bouten, W.; Camphuysen, K.C.J.; Nolet, B.A.; Shamoun-Baranes, J. (2022). Acceleration as a proxy for energy expenditure in a facultative‐soaring bird: comparing dynamic body acceleration and time‐energy budgets to heart rate. Funct. Ecol. 36(7): 1627-1638.
    Camphuysen, C.J. (2022). Seabirds and other charismatic megafauna in offshore habitats off NW Africa: GTA Ecological Vulnerability Analysis. NIOZ-rapport, 2022(03). NIOZ Royal Netherlands Institute for Sea Research: Texel. 80 pp.
    Camphuysen, C.J. (2022). Vulnerable wildlife concentrations at the Mauritanian Shelf - Atlas of area sensitivity to surface pollutants. NIOZ-rapport, 2022(04). NIOZ Royal Netherlands Institute for Sea Research: Texel. 48 pp.
    Camphuysen, C.J. (2022). Mission accomplished: chronic North Sea oil pollution now at acceptable levels, with Common Guillemots as sentinels. Seabird (2008) 34(1): 1-32
    Camphuysen, C.J.; Gear, S.C. (2022). Great Skuas and Northern Gannets on Foula, summer 2022 - an unprecedented, H5N1 related massacre. NIOZ-rapport, 2022(02). NIOZ Royal Netherlands Institute for Sea Research: Texel. 66 pp.
    Kentie, R.; Shamoun-Baranes, J.; Spaans, A.L.; Camphuysen, C.J. (2023). Spatial patterns in age‐ and colony‐specific survival in a long‐lived seabird across 14 contrasting colonies. Ibis 165(1): 82-95.
    Sage, E.; Bouten, W.; van Dijk, W.; Camphuysen, C.J.; Shamoun-Baranes, J. (2022). Built up areas in a wet landscape are stepping stones for soaring flight in a seabird. Sci. Total Environ. 852: 157879.
  • 2021
    Brown, J.M.; van Loon, E.E.; Bouten, W.; Camphuysen, C.J.; Lens, L.; Müller, W.; Thaxter, C.B.; Shamoun-Baranes, J. (2021). Long‐distance migrants vary migratory behaviour as much as short‐distance migrants: An individual‐level comparison from a seabird species with diverse migration strategies. J. Anim. Ecol. 90(5): 1058-1070.
    Camphuysen, C.J. (2021). Beached bird surveys in The Netherlands winter 2020/21. RWS Centrale Informatievoorziening BM 21.16. NIOZ Royal Netherlands Institute for Sea Research: Den Burg. 27 pp.
    Camphuysen, C.J. (2021). Vulnerable wildlife concentrations at the Mauritanian Shelf. A preliminary atlas of vulnerability to surface pollutants. NIOZ Royal Netherlands Institute for Sea Research: Den Burg. 46 pp.
    Camphuysen, C.J.; Kentie, R. (2021). Broedpoging van Geelpootmeeuw Larus michahellis op Texel in 2021. NIOZ Royal Netherlands Institute for Sea Research: Den Burg. 4 pp.
    Foekema, E.; van der Molen, J.; Asjes, A.; Bijleveld, A.; Brasseur, S.; Camphuysen, K. (C.J.); Van Franeker, J.A.; Holthuijsen, S.; Kentie, R.; Kühn, S.; Leopold, M.; Kleine Schaars, L.; Lok, T.; Niemann, H.; Schop, J. (2021). Ecologische effecten van het incident met de MSC Zoe op het Nederlandse Waddengebied, met focus op microplastics. NIOZ-rapport, 2021(03). NIOZ Royal Netherlands Institute for Sea Research: Texel. 98 pp.
    IJsseldijk, L.L.; Camphuysen, C.J.; Keijl, G.O.; Troost, G.; Aarts, G. (2021). Predicting harbor porpoise strandings based on near-shore sightings indicates elevated temporal mortality rates. Front. Mar. Sci. 8: 668038.
    Martín-Vélez, V.; Leeuwen, C.H.A.; Sánchez, M.I.; Hortas, F.; Shamoun-Baranes, J.; Thaxter, C.B.; Lens, L.; Camphuysen, C.J.; Green, A. (2021). Spatial patterns of weed dispersal by wintering gulls within and beyond an agricultural landscape. J. Ecol. 109(4): 1947-1958.
    Mercker, M.; Dierschke, V.; Camphuysen, C.J.; Kreutle, A.; Markones, N.; Vanermen, N.; Garthe, S. (2021). An indicator for assessing the status of marine-bird habitats affected by multiple human activities: A novel statistical approach. Ecol. Indic. 130: 108036.
    van Erp, J.A.; van Loon, E.E.; Camphuysen, C.J.; Shamoun-Baranes, J. (2021). Temporal patterns in offshore bird abundance during the breeding season at the Dutch North Sea coast. Mar. Biol. (Berl.) 168(10): 150.
  • 2020
    Bom, R.A.; van de Water, M.; Camphuysen, C.J.; van der Veer, H.W.; van Leeuwen, A. (2020). The historical ecology and demise of the iconic Angelshark Squatina squatina in the southern North Sea. Mar. Biol. (Berl.) 167(7): 91.
    Camphuysen, C.J. (2020). Dwarf eggs of herring gulls larus argentatus and lesser blackbacked gulls l. fuscus at texel, 2006-2020. Sula 28: 1-10
    van der Zwet, Z.; Camphuysen, K.; van der Meer, M.; Kentie, R. (2020). Van kuikenbloed naar kuikendieet. Tussen Dijk en Duin 1: 7-10
    Waggitt, J.; Peter G. H. Evans; Joana Andrade; Alex N. Banks; Oliver Boisseau; Mark Bolton; Gareth Bradbury; Tom Brereton; Cornelis Jan Camphuysen; Jan Durinck ; Tom Felce ; Ruben Christiaan Fijn ; Isabel Garcia‐Baron; Stefan Garthe; Steve C. V. Geelhoed; Anita Gilles; Martin Goodall; Jan Haelters; Sally Hamilton; Lauren Hartny‐Mills; Nicola Hodgins ; Kathy James; Mark Jessop ; Ailbhe S. Kavanag; Mardik Leopol; Katrin Lohrengel; Maite Louzao; Nele Markones; Jose Martínez-Cedeira; Oliver Ó Cadhla; Sarah L. Perr ; Graham J. Pierc ; Vincent Ridou ; Kevin P. Robinson; M. Begoña Santos; Camilo Saavedra; Henrik Skov; Eric W. M. Stienen; Signe Sveegaar; Paul Thompson; Nicolas Vanerme; Dave Wall; Andy Webb; Jared Wilson; Sarah Wanless; Jan Geert Hiddink (2020). Distribution maps of cetacean and seabird populations in the North‐East Atlantic. J. Appl. Ecol. 57(2): 253-269.
  • 2019
    Camphuysen, C.J. (2019). De onvoldoende begrepen dynamiek en samenhang van de Noordzee, in: Duimel, E. et al. (Ed.) Het roer moet om : Signalen voor een juiste koers van nieuw Noordzeebeleid. pp. 23-32
    Camphuysen, C.J. (2019). A decline in oil rates consolidated : monitoring and assessment of the proportion of oiled Common Guillemots in The Netherlands winter 2018/19. NIOZ Royal Netherlands Institute for Sea Research: Texel. 22 pp.
    Martín-Vélez, V.; Sánchez, M.I.; Shamoun-Baranes, J.; Thaxter, C.B.; Stienen, E.W.M.; Camphuysen, K.C.J.; Green, A.J. (2019). Quantifying nutrient inputs by gulls to a fluctuating lake, aided by movement ecology methods. Freshwat. Biol. 64(10): 1821-1832.
    Sage, E.; Bouten, W.; Hoekstra, B.; Camphuysen, K.C.J.; Shamoun-Baranes, J. (2019). Orographic lift shapes flight routes of gulls in virtually flat landscapes. NPG Scientific Reports 9(1).
    van Donk, S.; Dekker, R.; Dijkstra, A.; Drent, J.; Camphuysen, C.J. (2019). Eindrapportage monitoring mitigerende maatregelen Zwakke Schakels 2015-2018. NIOZ-rapport, 2019(2). NIOZ: Texel. 39 pp.
    van Donk, S.; Shamoun-Baranes, J.; Bouten, W.; van der Meer, J.; Camphuysen, C.J. (2020). Individual differences in foraging site fidelity are not related to time-activity budgets in Herring Gulls. Ibis 162(2): 429-445.
    van Donk, S.; Shamoun-Baranes, J.; Van der Meer, J.; Camphuysen, K.C.J. (2019). Foraging for high caloric anthropogenic prey is energetically costly. Movement Ecology 7(1).
  • 2018
    Bom, R.A.; Philippart, C.J.M.; van der Heide, T.; de Fouw, J; Camphuysen, C.J.; Dethmer, K.; Folmer, E.O.; Stocchi, P.; Stuut, J.-B.; van der Veer, H.W.; Al Zakwani, I. (2018). Barr Al Hikman: a pristine coastal ecosystem in the Sultanate of Oman: Current state of knowledge and future research challenges. NIOZ-rapport, 2018(1). NIOZ Royal Institute for Sea Research: Texel. 61 pp.
    Jouta, J.; de Goeij, P.; Lok, T.; Velilla, E.; Camphuysen, C.J.; Leopold, M.; van der Veer, H.W.; Olff, H.; Overdijk, O.; Piersma, T. (2018). Unexpected dietary preferences of Eurasian Spoonbills in the Dutch Wadden Sea: spoonbills mainly feed on small fish not shrimp. J. Ornithol. 159(3): 839-849.
    Keogan, K.; Daunt, F.; Wanless, S.; Phillips, R.A.; Walling, C.A.; Agnew, P.; Ainley, D.G.; Anker-Nilssen, T.; Ballard, G.; Barrett, R.T.; Barton, K.J.; Bech, C.; Becker, P.H.; Berglund, P.-A.; Bollache, L.; Bond, A.L.; Bouwhuis, S.; Bradley, R.W.; Burr, Z.M.; Camphuysen, C.J.; Catry, P.; Chiaradia, A.; Christensen-Dalsgaard, S.; Cuthbert, R.; Dehnhard, N.; Déscamps, S.; Diamond, T.; Divoky, G.; Drummond, H.; Dugger, K.M.; Dunn, M.J.; Emmerson, L.; Erikstad, K.E.; Fort, J.; Fraser, W.R.; Genovart, M.; Gilg, O.; González-Solis, J.; Granadeiro, J.P.; Gremillet, D.; Hansen, J.; Hanssen, S.A.; Harris, M.; Hedd, A.; Hinke, J.; Igual, J.M.; Jahncke, J.; Jones, I.; Kappes, P.J.; Lang, J.; Langset, M.; Lescroël, A.; Lorentsen, S.-H.; Lyver, P.O.; Mallory, M.; Moe, B.; Montevecchi, W.A.; Monticelli, D.; Mostello, C.; Newell, M.; Nicholson, L.; Nisbet, I.; Olsson, O.; Oro, D.; Pattison, V.; Poisbleau, M.; Pyk, T.; Quintana, F.; Ramos, J.A.; Ramos, R.; Reiertsen, T.K.; Rodríguez, C.; Ryan, P.; Sanz-Aguilar, A.; Schmidt, N.M.; Shannon, P.; Sittler, B.; Southwell, C.; Surman, C.; Svagelj, W.S.; Trivelpiece, W.Z.; Warzybok, P.; Watanuki, Y.; Weimerskirch, H.; Wilson, P.R.; Wood, A.G.; Phillimore, A.B.; Lewis, S. (2018). Global phenological insensitivity to shifting ocean temperatures among seabirds. Nat. Clim. Chang. 8(4): 313-318.
    Rakhimberdiev, E.; Duijns, S.; Karagicheva, J.; Camphuysen, C.J.; VRS Castricum; Dekinga, A.; Dekker, R.; Gavrilov, A.; ten Horn, J.; Jukema, J.; Saveliev, A.; Soloviev, M.Y.; Tibbitts, T.L.; van Gils, J.A.; Piersma, T. (2018). Fuelling conditions at staging sites can mitigate Arctic warming effects in a migratory bird. Nature Comm. 9: 4263.
    van Bemmelen, R.S.A.; Clarke, R.H.; Pyle, P.; Camphuysen, C.J. (2018). Timing and duration of primary molt in Northern Hemisphere skuas and jaegers. The Auk 135(4): 1043-1054.
  • 2017
    Camphuysen, C.J. (2017). Monitoring and assessment of the proportion of oiled Common Guillemots from beached bird surveys in The Netherlands: annual update winter 2015/16. NIOZ-rapport, 2017(1). Royal Netherlands Institute for Sea Research (NIOZ): Texel. 25 pp.
    Camphuysen, C.J.; van Bemmelen, R.; Van Spanje, T. (2017). Ship-based seabird and marine mammal surveys off Mauritania, 1-12 November 2016. Royal Netherlands Institute for Sea Research (NIOZ): Texel. 58 pp.
    Camphuysen, K. (C.J.); Henderson, P.A.; Velilla, E.; Kühn, S.; Leopold, M.F.; Somes, R. (2017). North Sea fish and their remains. Royal Netherlands Institute for Sea Research/Pisces Conservation Ltd: Texel. ISBN 978-1-90-4690-65-8. 326 pp.
    Goedknegt, M.A.; Havermans, J.; Waser, A.M.; Luttikhuizen, P.C.; Velilla, E.; Camphuysen, C.J.; Van der Meer, J.; Thieltges, D.W. (2017). Cross-species comparison of parasite richness, prevalence, and intensity in a native compared to two invasive brachyuran crabs. Aquat. Invasions 12(2): 201-212.
    Kompanje, E.J.O.; Camphuysen, C.J.; Leopold, M.F. (2017). The first case of conjoined twin harbour porpoises Phocoena phocoena (Mammalia, Cetacea). Deinsea-Rotterdam 17: 1-5
    Schulz, M.; Fleet, D.M.; Camphuysen, K.C.J.; Schulze-Dieckhoff, M.; Laursen, K. (2017). Oil pollution and seabirds, in: Kloepper, S. et al. Wadden Sea Quality Status Report 2017.
    van Donk, S.; Camphuysen, K.C.J.; Shamoun-Baranes, J.; Van der Meer, J. (2017). The most common diet results in low reproduction in a generalist seabird. Ecol. Evol. 7(13): 4620-4629.
  • 2016
    Camphuysen, C.J.; Smeenk, C. (2016). Noordkaper - Bruinvis, in: Broekhuizen, S. et al. (Ed.) Atlas van de Nederlandse zoogdieren. pp. 302-374
    Goedknegt, M.A.; Feis, M.E.; Wegner, K.M.; Luttikhuizen, P.C.; Buschbaum, C.; Camphuysen, C.J.; Van der Meer, J.; Thieltges, D.W. (2016). Parasites and marine invasions: Ecological and evolutionary perspectives. J. Sea Res. 113: 11–27.
    Rock, P.; Camphuysen, C.J.; Shamoun-Baranes, J.; Ross-Smith, V.; Vaughan, I.P. (2016). Results from the first GPS tracking of roof-nesting Herring Gulls Larus argentatus in the UK. Ringing & Migration 31(1): 47-62.
    Shamoun-Baranes, J.; Bouten, W.; van Loon, E.E.; Meijer, C.; Camphuysen, C.J. (2016). Flap or soar? How a flight generalist responds to its aerial environment. Philosophical Transactions of the Royal Society of London Series B-Biological Sciences 371(1704): 20150395.
  • 2015
    Camphuysen, C.J. (2015). Monitoring and assessment of the proportion of oiled Common Guillemots from beached bird surveys in The Netherlands: update winter 2014/15. NIOZ-rapport, 2015(4). NIOZ: Den Burg. 26 pp.
    Camphuysen, C.J.; Kloff, S.; Jiyid'ould Taleb, M.A. (2015). Ship-based seabird and marine mammal surveys off Mauritania, 4-14 September 2015. NIOZ-rapport, 2015(2). NIOZ: Den Burg. 102 pp.
    Camphuysen, K.C.J.; Shamoun-Baranes, J.; van Loon, E.E.; Bouten, W. (2015). Sexually distinct foraging strategies in an omnivorous seabird. Mar. Biol. (Berl.) 162(7): 1417-1428.
    Hoekendijk, J.P.A.; de Vries, J.J.; van der Bolt, K.; Greinert, J.; Brasseur, S.; Camphuysen, K.C.J.; Aarts, G. (2015). Estimating the spatial position of marine mammals based on digital camera recordings. Ecol. Evol. 5(3): 578–589.
    IJsseldijk, L.L.; Camphuysen, K.C.J.; Nauw, J.J.; Aarts, G. (2015). Going with the flow: Tidal influence on the occurrence of the harbour porpoise (Phocoena phocoena) in the Marsdiep area, The Netherlands. J. Sea Res. 103: 129-137.
    Tyson, C.; Shamoun-Baranes, J.; van Loon, E.E.; Camphuysen, K.C.J.; Hintzen, N.T. (2015). Individual specialization on fishery discards by lesser black-backed gulls (Larus fuscus). ICES J. Mar. Sci./J. Cons. int. Explor. Mer 72(6): 1882-1891.
  • 2014
    Camphuysen, C.J. (2014). Terug van weggeweest: Bruinvissen, in: Groen, N. et al. Texel is anders : landschap, (cultuur)historie en natuur. pp. 130-133
    Camphuysen, C.J. (2014). Zeezoogdieren, in: van der Mark, C. et al. Zeeboek: determinatietabellen voor flora en fauna van de Nederlandse kust. pp. 211-219
    van der Mark, C.; Ates, R.; De Boois, I.; Bor, P.; Buizer, B.; van der Burgt, F.; Camphuysen, K.; Dekker, W.; Driessen, F.; Den Hartog, K.; Huijsman, M.; Karremans, M.; Oosterbaan, A.; Otten, B.G.; Perk, F.; de Ruijter, E.; de Ruijter, G.; van Santbrink, J.; Schoenmaker, A.; van Soest, R.; Stegenga, H.; Tulp, A. (2014). Zeeboek: determinatietabellen voor flora en fauna van de Nederlandse kust. Stichting Jeugdbondsuitgeverij: 's Graveland. ISBN 978-90-5107-000-2. 240 pp.
  • 2013
    Bicknell, A.W.J.; Oro, D.; Camphuysen, K.; Votier, S.C. (2013). Potential consequences of discard reform for seabird communities. J. Appl. Ecol. 50(3): 649-658.
    Bouten, W.; Baaij, E.W.; Shamoun-Baranes, J.; Camphuysen, K.C.J. (2013). A flexible GPS tracking system for studying bird behaviour at multiple scales. J. Ornithol. 154(2): 571-580.
    Camphuysen, K.C.J. (2013). A historical ecology of two closely related gull species (Laridae) : multiple adaptations to a man-made environment. PhD Thesis. Rijksuniversiteit Groningen: [s.l.]. ISBN 9789090275383. 410 pp.
    Cox, S.L.; Scott, B.E.; Camphuysen, C.J. (2013). Combined spatial and tidal processes identify links between pelagic prey species and seabirds. Mar. Ecol. Prog. Ser. 479: 203-221.
    Peltier, H.; Baagøe, H.J.; Camphuysen, K.C.J.; Czeck, R.; Dabin, W.; Daniel, P.; Deaville, R.; Haelters, J.; Jauniaux, T.; Jensen, L.F.; Jepson, P.D.; Keijl, G.O.; Siebert, U.; Van Canneyt, O.; Ridoux, V. (2013). The stranding anomaly as population indicator: the case of Harbour Porpoise Phocoena phocoena in North-Western Europe. PLoS One 8(4): 1-14.
  • 2012
    Camphuysen, C.J.; Gronert, A. (2012). Apparent survival and fecundity of sympatric Lesser Black-backed Gulls and Herring Gulls with contrasting population trends. Ardea 100(2): 113-122.
    Camphuysen, C.J.; Shamoun-Baranes, J.; Bouten, W.; Garthe, S. (2012). Identifying ecologically important marine areas for seabirds using behavioural information in combination with distribution patterns. Biol. Conserv. 156: 22-29.
    Embling, C.B.; Illian, J.; Armstrong, E.; van der Kooij, J.; Sharples, J.; Camphuysen, K.C.J.; Scott, B.E. (2012). Investigating fine-scale spatio-temporal predator-prey patterns in dynamic marine ecosystems: a functional data analysis approach. J. Appl. Ecol. 49(2): 481-492.
    Hallgrimsson, G.T.; Gunnarsson, H.V.; Torfason, O.; Buijs, R.-J.; Camphuysen, K.C.J. (2012). Migration pattern of Icelandic Lesser Black-backed Gulls Larus fuscus graellsii: indications of a leap-frog system. J. Ornithol. 153(3): 603-609.
    Shamoun-Baranes, J.; van Loon, E.E.; Purves, R.S.; Speckmann, B.; Weiskopf, D.; Camphuysen, C.J. (2012). Analysis and visualization of animal movement. Biol. Lett. 8: 10.1098/rsbl.2011.0764 .
  • 2011
    Bartolomé, C.; Maside, X.; Camphuysen, C.J.; Heubeck, M.; Bao, R. (2011). Multilocus population analysis of Gavia immer (Aves: Gaviidae) mtDNA reveals low genetic diversity and lack of differentiation across the species breeding range. Org. Divers. Evol. 11(4): 307-316.
    Camphuysen, C.J. (2011). Lesser Black-backed Gulls nesting at Texel: Foraging distribution, diet, survival, recruitment and breeding biology of birds carrying advanced GPS loggers. NIOZ-rapport, 2011(5). NIOZ: Texel. 80 pp.
    Camphuysen, C.J.; de Leeuw, J.W. (2011). Environmental impact of hydrophobic chemicals with low water solubility released from merchant shipping, in: Roose, P. et al. Monitoring Chemical Pollution in Europe’s Seas: Programmes, practices and priorities for research. European Marine Board Position Paper, 16: pp. 51-63
    Camphuysen, C.J.; Vercruijsse, H. J. P. ; Spaans, A.L. (2011). Colony- and age-specific seasonal dispersal of Herring Gulls Larus argentatus breeding in The Netherlands. J. Ornithol. 152(4): 849-868.
    Camphuysen, K.C.J. (2011). Seabirds and chronic oil pollution: Self-cleaning properties of gulls, Laridae, as revealed from colour-ring sightings. Mar. Pollut. Bull. 62(3): 514-519.
    Camphuysen, K.C.J. (2011). Kannibalisme bij de Kleine Mantelmeeuw Larus fuscus op Texel: een gedocumenteerd geval Cannibalism in Lesser Black-Backed Gulls at Texel, a Documented Case. Sula 24(2): 83-90
    Camphuysen, K.C.J. (2011). Olieslachtoffers op de Nederlandse Kust, 2010/2011 = Oiled Seabirds Washing Ashore in the Netherlands, 2010/2011. NIOZ: Texel. 24 pp.
    Camphuysen, K.C.J. (2011). Northern Gannets in the North Sea: foraging distribution and feeding techniques around the Bass Rock. Br. Birds 104: 60-76
    Camphuysen, K.C.J.; Gronert, A. (2011). Gekleurringde, plaatstrouwe Zilvermeeuwen Larus argentatus op strandhoofden in Noord-Holland verliezen voedselbron? = Colour-Ringed, Site-Faithful Herring Gulls at Breakwaters in Noord-Holland Lose Resource?. Sula 24(2): 61-74
    Camphuysen, K.C.J.; Krop, A. (2011). Maternal care, calf-training and site fidelity in a wild harbour porpoise in the North Sea. Lutra 54(2): 123-126
    Camphuysen, K.C.J.; Siemensma, M.L. (2011). Conservation plan for the Harbour Porpoise Phocoena phocoena in The Netherlands: towards a favourable conservation status. NIOZ-rapport, 2011(7). NIOZ: Texel. 183 pp.
    Camphuysen, K.C.J.; van Bemmelen, R. (2011). Herkenning van Vorkstaartmeeuwen Xema sabini in de hand op het strand = Identification of Sabine’s Gulls in the Hand during Beached Bird Surveys. Sula 24(2): 49-60
    Roose, P.; Albaigés, J.; Bebianno, M.J.; Camphuysen, C.J.; Cronin, M.; de Leeuw, J.; Gabrielsen, G.; Hutchinson, T.; Hylland, K.; Jansson, B.; Jenssen, B.M.; Schulz-Bull, D.; Szefer, P.; Webster, L.; Bakke, T.; Janssen, C.; Calewaert, J.-B.; McDonough, N. (2011). Monitoring Chemical Pollution in Europe’s Seas: Programmes, practices and priorities for research. European Marine Board Position Paper, 16. Marine Board-ESF: Ostend. ISBN 978-2-918428-57-2. 103 pp.
    Shamoun-Baranes, J.; Bouten, W.; Camphuysen, C.J.; Baaij, E. (2011). Riding the tide: intriguing observations of gulls resting at sea during breeding. Ibis 153(2): 411-415.
    Wojczulanis-Jakubas, K.; Jakubas, D; Welcker, J.; Harding, A.M.A.; Karnovsky, N.J.; Kidawa, D.; Steen, H.; Stempniewicz, L.; Camphuysen, C.J. (2011). Body size variation of a high-Arctic seabird: the dovekie (Alle alle). Polar Biol. 34(6): 847-854.

Linked projects

Effects of invasive species
David Thieltges
Netherlands Organization for Scientific Research - Ocean and Coastal Research
Project duration
31 Dec 2011 - 30 Aug 2020
Diet of Gulls
Myron Peck
Netherlands Organization for Scientific Research - Dynamisering
Project duration
1 Jun 2013 - 31 Dec 2018
UUNIOZ_The role of the aviant nutrient pump for back-barrier island dynamics
Kees Camphuijsen
Utrecht University
Project duration
1 Jan 2021 - 31 Dec 2025
ACTNOW_Advancing understanding of Cumulative Impacts on European marine biodiversity
Myron Peck
European Community
Project duration
1 Jan 2023 - 31 Dec 2026
Monitoring beached bird surveys - measuring the effects of chronic oil pollution
Kees Camphuijsen
Project duration
1 Jan 2017 - 31 Dec 2020
UUNIOZ_The role of the aviant nutrient pump for back-barrier island dynamics
Kees Camphuijsen
Utrecht University
Project duration
1 Jan 2021 - 31 Dec 2025
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