Kees (C.J.) Camphuysen

Marine ornithologist

Expertise

Marine Ecology - Foraging ecology of seabirds - Marine mammals - Tracking studies of seabirds - Colony studies - Offshore surveys

Research interests

CV

 Professional career

Associated with the Royal Netherlands Institute for Sea Research (Royal NIOZ) since 1992, first employed on a research project on the effect of beamtrawl fisheries on seabirds^17-20. Co-ordinated two EC funded projects, on behalf of NIOZ studying the effects of discards on seabirds on a North Sea scale (Discards I and Discards II)^21-25. Later work focused mainly on natural aspects underlying the distribution of seabirds at sea^25-27, which culminated in the invitation to join another EC funded project in which a complicated model of foraging decisions of seabirds was parameterised and tested (MIFOS project)^{28,47,http://www.abdn.ac.uk/mifos/}. Initiated and co-ordinated a further EC funded research project (IMPRESS, 2000-2004) ^{ttp://www.nioz.nl/en/deps/mee/projects/impress/impress.htm} evaluating the effects of industrial sandeel fisheries on seabirds. This project was finalised in 2005. The work on fisheries impacts has culminated in several topical papers^{40,43-44,48,50-52,59}, a book^{Cambridge University Press}, book chapters, and numerous reports^{see publications, 2005}.

Formed consultancy in 1995 (CSR Consultancy), closely associated with Royal NIOZ and IBN-DLO/Alterra (currently Wageningen IMARES) in which applied scientific questions are addressed, such as environmental impact assessments in the North Sea and Wadden Sea for governmental bodies, NGOs and oil companies. Consultancy discontinued in 2006 because of a permanent research position at Royal NIOZ.

Studies of the distribution of seabirds at sea (mainly 1985-present) have culminated in the participation in the publication of a number of atlases, books and topical papers^{29-31,39,45,53,58}. Studies on cetaceans commenced in 1982 and have culminated into several publications on the identification and distribution of marine mammals^{12, 32-38} (1982-present). Recent work is mainly aiming at the integration of studies of important top-predators in marine ecosystems, mainly seabirds and cetaceans^39-42. A Marine mammal database was established in 1987 and has been maintained as a database manager for the Dutch Seabird group and this database is now the primary and most accessible source of information for the occurrence and relative abundance of cetaceans in the southern North Sea. Personal experience with 50 taxa of cetaceans; personal records: 25,000 individual whales and dolphins (as in 2012). Author of two field guides, one published in Norwegian, one in Dutch, and several papers on the identification of whales and dolphins. Author of a book on whales and dolphins in the North Sea (2006). Conducted mass autopsies of Harbour Porpoises stranded in The Netherlands in 2006 and 2007, 2008, and 2009 and published the results as a report (NIOZ/IMARES project)^{see reports under Publications}and in a topical paper on marine mammals strandings in The Netherlands⁶⁵.

Participated in two and led or guided 12 expeditions into the arctic between 1982 and 2009 (Greenland, Jan Mayen, Greenland Sea, Iceland, Svalbard), which were purely for scientific research (5x) or as tourist guide (10x). Participated in three scientific cruises with TNO-FEL in 2005-2006 in the Bay of Biscay, the Mediterranean, and the Lofoten area as whale expert during sightings surveys with acoustic arrays being operated simultaneously (acoustic trials to identify marine mammals that were logged with high tech acoustic equipment onboard Royal navy vessels Mercuur and Snellius and the Norwegian Forsvarets Forsknings Institut’s vessel Sverdrup II ^{see reports under ‘Publications’}.

            Operated as tourist guide in the Antarctic in 2007, 2008, 2009, and 2010. Studies included seabird population censuses^10-12,seabird migration studies¹³, diet studies^14,15, and systematic surveys for the distribution of seabirds and cetaceans¹⁶. Participated in scientific cruises studying the offshore foraging ecology of seabirds and marine mammals and distribution patterns in relation with oceanographical conditions in the South Atlantic in 2000^46,61 and 2001^56-57 under contract with the Netherlands Institute for Sea Research (RV Pelagia), and to South Georgian waters in 2002 under contract with the British Antarctic Survey (BAS, Cambridge, UK; RV James Clark Ross)⁶³. Contracted for studies on the abundance and distribution of seabirds and marine mammals off Mauritania in 2003 by Woodside Energy Ltd (Australia) and in 2012 by GIZ/GEF/IMROP.

            Scientific studies under the beached bird survey project include studies of morphometrics, age composition, sex ratio and physical condition of wrecked seabirds, the same for birds involved in major oil spills (Amoco Cadiz, Prestige, Tricolor, most recently in a spill in Estonia in 2006 and a spill in The Netherlands in 2007), a skull ossification project with Edward Soldaat (examining ageing characteristics in auks) and a study of post-nuptial moult and chick development in [stranded] Common Guillemots Uria aalge. Ongoing studies of the diet of auks based on stranded specimens.

            Co-organiser of the World Seabird Group Conference, Victoria (2010). Elected board member (European representative) in World Seabird Union (2013- )

             In 2006, I launched a new project from Royal NIOZ, to evaluate the reproductive success and feeding ecology of Herring Gulls Larus argentatus and Lesser Black-backed Gulls Larus fuscus, including breeding in the Dutch Wadden Sea, to evaluate the effects of prey availability fluctuations, individual levels of specialisation and habitat use, and other factors that might explain the recent failures and population declines in the Herring Gull and the consistently high levels of reproduction and population increase in the Lesser Black-backed Gull. A colour-ring programme was developed and set-up, which is the successor of the Alterra/Arie Spaans colour ringing programme that was transferred from Alterra to Royal NIOZ in 2005. An analysis of the dispersal and annual survival of Herring Gulls ringed between 1986 and 1988 in 14 Dutch colonies has been published⁶⁶. Radio telemetry studies in the gull colonies were conducted to evaluate levels of parental care. A large diet study was conducted to evaluate shifts in prey utilised throughout the breeding season and to study individual specialisation. The first papers on these gull studies have meanwhile been published^{Limosa & Seevögel, see Publications}. In 2008 field experiments were conducted to evaluate the possibilities of nest attendance enhancement through supplementary food (RUG masterproject). In a collaboration with University of Amsterdam (Prof W Bouten, D J Shamoun-Baranes), cutting edge technology GPS loggers devices have been deployed in 2008-2012, to evaluate foraging distribution patterns of individual birds in great detail^67-69.

Qualifications for offshore work: Participated in >100 offshore trips on board a variety of (research) vessels (ca. 2-3 months per annum in recent years, up to 5 months per annum in earlier years), gas production platforms (1985/85, 2005) and at Meetpost Noordwijk (offshore platform; for approx. 15 weeks 1978-82). Completed basic offshore safety and emergency response training, including helicopter under water escape techniques (HUET) and airpocket (Nogepa 0.5A), completed safety courses for windfarm-related offshore studies.

Ship-based surveys: around 100,000 km steamed in Northern Hemisphere (Mediterranean Sea, Bay of Biscay, North Sea, Channel, west of Britain, Irish Sea, Celtic Sea, Norwegian Sea, Greenland Sea, Davis Strait, Denmark Strait, Barentsz Sea), 10,400 km steamed Africa and Southern Hemisphere (Northwest Africa, South Africa, Namibia, Uruguay - South Georgia – Falklands, South Orkneys, Antarctic Peninsula). International trainer of European Seabirds at Sea database programme for ship-based surveys (at-sea methods) and data analysis (data processing and database management).

Qualifications for colony work: Co-organiser and performer of colony censuses at Jan Mayen in 1983 (all species, including tundra-nesting birds). Participated in colony censuses for Black Guillemot (Western Isles, with M.L. Tasker), Atlantic Puffin (St Kilda, with M.P. Harris and M.L. Tasker), and ‘seabird colonies’ (multispecies; Scottish east coast, with M.L. Tasker).

Established long-term monitoring programme for colonial Herring Gulls and Lesser Black-backed Gulls in 2006 at texel (The Netherlands), recording breeding success with the help of enclosures, diet studies (pellets and regurgitated material), recording nest attendance (radio telemetry), recording foraging distribution (using advanced GPS loggers, collaboration with the University of Amsterdam), recording and analysing chick growth, establishing and managing a colour-ring programme for demographic studies at the Royal NIOZ.

Autopsies stranded seabirds: Organiser and performer thousands of autopsies of (oiled) stranded seabirds, including diet studies of seabirds from stomach contents and regurgitated material. Methods recently described and standardised in Handbook Oil Impact Assessment⁶⁴. Performed mass necropsies in association with oil spills around Europe.

Seabird diet studies: Studies of seabird diets have ranged from high-arctic Northern Fulmars to southern European wintering seabirds including Great Northern Divers, various species of auks and other seabirds. Within The Netherlands, emphasis of dietary studies on Alcidae and large gulls. Extensive experience with the identification of fish bones and otoliths, considerable experience with the identification of undigestable remains of mammals and birds, human refuse and other prey items. Manager of NIOZ seabird diet reference collection.

Qualifications as a seabird ringer: Ringing seabirds under Norwegian license at Jan Mayen in 1983 (Atlantic Puffin, Common Guillemot, Little Auk, Brünnich’s Guillemot), ringing Northern Fulmars under Norwegian licence at Svalbard (several years, 1980s), ringing and colour-ringing Herring Gulls and Lesser black-backed Gulls in The Netherlands under Dutch licence (2006-present). Colour-ring database manager Spaans/NIOZ database, member of advisory board of the Dutch ringing centre.

Qualifications for animal experiments: Ontheffing artikel 12 van de wet op de dierproeven (9 June 2010) krachtens artikel 16 Wod tweede lid.

Data loggers: Extensive experience with the successful deployments of dataloggers (with GPS) and radio tags (2006-2012) on Herring Gulls and Lesser black-backed Gulls.

Computer skills: standard but long-term experience with word processing software, spreadsheet use, and several statistical packages (25 years experience). Extensive experience with database design and database management. Experience with several mapping programmes and statistical package r.

Editorships: Editor of Sula (journal of the Dutch Seabird group) during 1987-1998 and again in 2008-present. (Technical) editor of Ardea (scientific journal Netherlands Ornithologists Union) 1996-2005, founder and co-editor of Atlantic Seabirds 1999-2007 (scientific journal of two seabird groups in Europe), member editorial board of Marine Ornithology 2000-2010, member editorial board of Lutra(Journal of Mammalogy, The Netherlands) during 2007-2012. Member editorial board of Seabird since 2008 (international journal of the Seabird Group, UK). Currently member editorial board ICES Journal of Marine Science (2013- ).

Teaching: Guest lecturer University Leiden (2007, 2008) and University Utrecht (2010, 2011, 2012) on marine mammals in Marine Sciences courses.

Awards Received Herman Klomp award in 1993, for 150 publications written as an amateur to that date, but particularly for two papers on seabirds and fisheries interactions^{3, 17}, from NOU, SOVON and Vogelbescherming Nederland.

Publications specifically mentioned in CV

¹Camphuysen C.J. & Dijk J.van 1983. Zee- en kustvogels langs de Nederlandse kust, 1974-79. Limosa 56: 81-230.

²Camphuysen C.J. 1989. Beached Bird Surveys in the Netherlands 1915-1988; Seabird Mortality in the southern North Sea since the early days of Oil Pollution. Techn. Rapport Vogelbescherming 1, Werkgroep Noordzee, Amsterdam 322pp.

³Camphuysen C.J. 1992. Zeevogelstrandingen op de Nederlandse kust: 26 jaar een vinger aan de pols (1965-1991). Limosa 66(1): 1-16.

⁴Camphuysen C.J. 1998. Beached bird surveys indicate decline in chronic oil pollution in the North Sea. Mar. Poll. Bull. 36: 519-526.

⁵Camphuysen C.J. 1995. Olieslachtoffers langs de Nederlandse kust als indicatoren van de vervuiling van de zee met olie. Sula 9(special issue): 1-90, I-XX.

⁶Camphuysen C.J. 1997. Olievervuiling en olieslachtoffers langs de Nederlandse kust, 1969-97: signalen van een schonere zee. Sula 11(special issue): 41-156.

⁷ Camphuysen C.J. & Dahlmann G. 1995. Guidelines on standard methodology for the use of (oiled) beached birds as indicators of marine pollution. Ad Hoc working group on Monitoring, Oslo and Paris Convention for the Prevention of Marine Pollution. MON 95/7, Agenda item 7, 13-17 November 1995, Copenhagen.

⁸Camphuysen C.J. & Franeker J.A. van 1992. The value of beached bird surveys in monitoring marine oil pollution. Techn. Rapport Vogelbescherming 10, Vogelbescherming Nederland, Zeist, 191pp.

⁹Dahlmann G., Timm D., Averbeck C., Camphuysen C.J. & Skov H. 1994. Oiled seabirds - Comparative investigations on oiled seabirds and oiled beaches in the Netherlands, Denmark and Germany (1990-1993). Mar. Poll. Bull. 28: 305-310.

¹⁰Franeker J.A. van, Camphuysen C.J. & Mehlum F. 1986. Status over Jan Mayens fugler. Vår Fuglefauna 9: 145-158.

¹¹Franeker J.A. van, Camphuysen C.J. & Mehlum F. 1998. The birds of Jan Mayen. Circumpolar Journal 13(2): 28-43.

¹²Camphuysen C.J. 1993. Birds and (marine) mammals in Svalbard, 1985-91. Sula 7(special issue): 3-44.

¹³Camphuysen C.J. 1989. Diurnal rhythm of the Fulmar Fulmarus glacialis in the arctic summer. Dansk Orn. Foren. Tidsskr. 83: 85-86.

¹⁴Camphuysen C.J. & van Franeker J.A. 1988. Diet of the Fulmar Fulmarus glacialis on Bear Island, Jan Mayen and Spitsbergen. In: Tasker M.L. (ed.). Seabird Food and Feeding Ecology: 14-15. Proc. 3rd Int. Conf. Seabird Group, Cambridge, 12-14 Febr. 1988.

¹⁵Camphuysen C.J. & Franeker J.A. van 1997. Notes on the diet of Northern Fulmars Fulmarus glacialis from Bjørnøya (Bear Island). Sula 11: 1-10.

¹⁶Camphuysen C.J. 1993. Summer distribution of seabirds and marine mammals in the Greenland Sea, 1985-90. Sula 7(special issue): 45-64.

¹⁷Camphuysen C.J. 1993. Foerageermogelijkheden voor zeevogels in de boomkorvisserij: een verkennend onderzoek. Sula 7: 81-104.

¹⁸Camphuysen C.J. 1994. Flatfish selection by Herring Gulls Larus argentatus and Lesser Black-backed Gulls Larus fuscus scavenging at commercial beamtrawlers in the southern North Sea. Neth. J. Sea Res. 32: 91-98.

¹⁹Camphuysen C.J. 1994. Scavenging seabirds at beamtrawlers in the southern North Sea: distribution, relative abundance, behaviour, prey selection, feeding efficiency, kleptoparasitism, and the possible effects of the establishment of 'protected areas'. BEON Report 1994-14, Netherlands Institute for Sea Research, Texel.

²⁰Camphuysen C.J. 1995. Herring Gull Larus argentatus and Lesser Black-backed Gulls Larus fuscus feeding at fishing vessels in the breeding season: competitive scavenging versus efficient flying. Ardea 83: 365-380.

²¹Camphuysen C.J., Ensor K., Furness R.W., Garthe S., Hüppop O., Leaper G., Offringa H. & Tasker M.L. 1993. Seabirds feeding on discards in winter in the North Sea. Final report to the European Comm., study contr. 92/3505, NIOZ-Report 1993-8, Netherlands Institute for Sea Research, Texel.

²²Camphuysen C.J., Calvo B., Durinck J., Ensor K., Follestad A., Furness R.W., Garthe S., Leaper G., Skov H., Tasker M.L. & Winter C.J.N. 1995. Consumption of discards by seabirds in the North Sea. Final report to the European Comm., study contr. BIOECO/93/10, NIOZ-Report 1995-5, Netherlands Institute for Sea Research, Texel, 202+LVIIIpp.

²³Garthe S., Camphuysen C.J. & Furness R.W. 1996. Amounts of discards in commercial fisheries and their significance as food for seabirds in the North Sea. Mar. Ecol. Progr. Ser. 136: 1-11.

²⁴Camphuysen C.J. 1995. Kittiwakes Rissa tridactyla in the North Sea: pelagic ecology, fisheries relationships and feeding strategies. In: Camphuysen C.J. (ed.). Proceedings NOU/ESAS symposium Seabirds at sea in the North Sea, Texel, 8 October 1994. Limosa 68: 123.

²⁵Camphuysen C.J. & Garthe S. 1997. Distribution and scavenging habits of Northern Fulmars in the North Sea. ICES J. Mar. Sc. 54: 654-683.

²⁶Camphuysen C.J. 1998. Het voorkomen van de Alk Alca torda in Nederlandse wateren. Limosa 71: 69-77.

²⁷Camphuysen C.J. 1999. Diurnal activity patterns and nocturnal group formation of wintering Common Murres in the central North Sea. Colonial Waterbirds 21: 406-413.

²⁸Camphuysen C.J. & A. Webb 1999. Multi-species feeding associations in North Sea seabirds: jointly exploiting a patchy environment. Ardea 87: 177-198.

²⁹Camphuysen C.J. & Leopold M.F. 1994. Atlas of seabirds in the southern North Sea. IBN Research report 94/6, NIOZ-Report 1994-8, Institute for Forestry and Nature Research, Netherlands Institute for Sea Research and Dutch Seabird Group, Texel.

³⁰Stone C.J., Webb A., Barton C., Ratcliffe N., Reed T.C., Tasker M.L., Camphuysen C.J. & Pienkowski M.W. 1995. An atlas of seabird distribution in north-west European waters. Joint Nature Conservation Committee, Peterborough, 326pp.

³¹Camphuysen C.J. 1996. De verspreiding van zeevogels in de Noordzee: naar een beter begrip van patronen en verbanden. Sula 10(special issue 2): 41-88.

³²Camphuysen C.J. 1987. Het ontdekken en herkennen van Bruinvissen Phocoena phocoena op zee. Sula 1(3): 66-72.

³³Camphuysen C.J. 1987. Handleiding voor de determinatie van walvisachtigen (Cetacea) in het Noordoost-Atlantische gebied. Uitgave eigen beheer, Zaandam, 2e druk (januari 1987) pp 1-34.

³⁴Camphuysen C.J. 1991. Hvalguide - Veiledning i bestemmelse av hval i de Nordøst-Atlantiske havområder. Norsk Inst. Naturforsk., Trondheim NINA Temahefte 1: 1-41.

³⁵Camphuysen C.J. & Ouden J.E. den 1988. Geassocieerd voorkomen van zeevogels en Dwergvinvissen Balaenoptera acutorostrata in de Noordzee, september 1988. Sula 2(3): 92-93.

³⁶Camphuysen C.J. & Leopold M.F. 1993. The Harbour Porpoise Phocoena phocoena in the southern North Sea, particularly the Dutch sector. Lutra 36(1): 1-24.

³⁷Camphuysen C.J. 1994. The Harbour Porpoise Phocoena phocoena in the southern North Sea, II: a come-back in Dutch coastal waters? Lutra 37(1): 54-61.

³⁸Northridge S., Tasker M.L., Webb A., Camphuysen C.J. & Leopold M.F. 1997. White-beaked Lagenorhynchus albirostris and Atlantic white-sided L. acutus distributions in north-west European and U.S. North Atlantic waters. SC/48/SM44, Rep. Int. whal. Commn. 47: 797-805.

³⁹Bourne W.R.P. & Camphuysen C.J. 2000. Seabirds. Chapter 114, In: Sheppard C. (ed.) Seas at the Millenium, an environmental evaluation: 105-116. Elsevier, Amsterdam.

⁴⁰Camphuysen C.J. 1999. New feeding technique of Great Cormorants Phalacrocorax carbo sinensis at beam trawlers. Atlantic Seabirds 1(2): 85-90.

⁴¹Camphuysen C.J., Wright P.J., Leopold M.F., Hüppop O. & Reid J.B. 1999. A review of the causes, and consequences at the population level, of mass mortalities of seabirds. In: Furness R.W. & Tasker M.L. (eds) Diets of seabirds and consequences of changes in food supply: 51-66. ICES Coop. Res. Report No. 232, International Council for the Exploration of the Sea, Copenhagen.

⁴²Camphuysen C.J. 2001. Seabirds and fisheries interaction. In: Steele J.H., Thorpe S.A. & Turekian K.K. (eds) Encyclopedia of Ocean Sciences: 2677-2686. Academic Press, London.

⁴³Tasker M.L., Camphuysen C.J., Cooper J., Garthe S., Montevecchi W.A. & Blaber S.J.M. 2000. The impacts of fishing on marine birds. ICES J. Mar. Sc. 57: 531-547.

⁴⁴Camphuysen C.J. & Garthe S. 2000. Seabirds and commercial fisheries: population trends of piscivorous seabirds explained? Chapter 11 In: Kaiser M.J. & Groot S.J. de (eds). Effects of fishing on non-target species and habitats: Biological, Conservation and Socio-Economic Issues: 163-184. Blackwell Science, Oxford.

⁴⁵Camphuysen C.J. 2000. Zomerwaarnemingen van Noordse Pijlstormvogels Puffinus puffinus in Nederland. Limosa 73: 7-16.

⁴⁶Camphuysen C.J. & J. van der Meer 2000. Notes on the distribution of the Spectacled Petrel Procellaria conspicillata in the South Atlantic. Atlantic Seabirds 2(1): 13-18.

⁴⁷Camphuysen C.J. & J. van der Meer 2000. The distribution of seabirds and their prey at the fine scale. section 3.2 In: Ollason J.G. (ed.) Modelling the Impact of Fisheries on Seabirds: 218-233. Final Report CFP 96-079, University of Aberdeen, Aberdeen.

⁴⁸Camphuysen C.J. & Hüppop O. 2000. Review of the extent to which fisheries have altered the composition of seabird communities. In: Tasker M.L. (ed.) Report of the Working Group on Seabird Ecology, Wilhelmshaven, Germany, 20-23 March 2000: 18-34. Oceanography Committee, ICES CM 2000/C:04, Ref. ACME+E, International Council for the Exploration of the Sea, Copenhagen, Denmark.

⁴⁹Camphuysen C.J. 2000. Further review of the contents on the database on seabird diet composition. In: Tasker M.L. (ed.) Report of the Working Group on Seabird Ecology, Wilhelmshaven, Germany, 20-23 March 2000: 54-57. Oceanography Committee, ICES CM 2000/C:04, Ref. ACME+E, International Council for the Exploration of the Sea, Copenhagen, Denmark.

⁵⁰Camphuysen C.J. 2000. Mass mortality of Common Eiders Somateria mollissima in the Wadden Sea, winter 1999/2000: food related parasite outbreak? Atlantic Seabirds 2(1): 47-48.

⁵¹Camphuysen C.J. 2000. Seabirds drowned in fishing nets off Jan Mayen (Greenland Sea). Atlantic Seabirds 2(2): 87-91.

⁵²Tasker M.L., Camphuysen C.J. & Garthe S. 2000. The spectacular spread of Northern Fulmars Fulmarus glacialis in the North Atlantic over the last 300 years: facts and fairy tales reviewed. In: Flint E. & Swift K. (eds) Second international conference on the biology and conservation of albatrosses and other petrels, Honolulu, Hawaii, USA, 8-12 May 2000; abstracts of oral and poster presentations. Mar. Ornithol. 28: 149.

⁵³Bijlsma R.G., Hustings F. & Camphuysen C.J. 2001. Schaarse en algemene vogels van Nederland. Avifauna van Nederland, 2. KNNV Uitgeverij Utrecht en GMB Uitgeverij, Haarlem, 497pp.

⁵⁴Camphuysen C.J. & Heubeck M. 2001. Marine oil pollution and beached bird surveys: the development of a sensitive monitoring instrument. Environmental Pollution 112: 443-461.

⁵⁵Camphuysen C.J. 2001. Northern Gannets Morus bassanus found dead in The Netherlands, 1970-2000. Atlantic Seabirds 3(1): 15-30.

⁵⁶Camphuysen C.J. & Van der Meer J. 2001. Pelagic distribution, moult and (sub-) specific status of Cory's Shearwaters Calonectris [d.] diomedea/borealis wintering off southern Africa. Marine Ornithology 29: 85-93.

⁵⁷Camphuysen C.J. 2001. The distribution of Spectacled Petrels Procellaria conspicillata in the south-eastern Atlantic. Atlantic Seabirds 3(3): 113-124.

⁵⁸Camphuysen C.J. 2002. Post-fledging dispersal of Common Guillemots Uria aalge guarding chicks in the North Sea: the effect of predator presence and prey availability at sea. Ardea 90(1): 103-119.

⁵⁹Camphuysen C.J., C.M. Berrevoets, H.J.W.M. Cremers, A. Dekinga, R. Dekker, B.J. Ens, T.M. van der Have, R.K.H. Kats, T. Kuiken, M.F. Leopold, J. van der Meer & T. Piersma 2002. Mass mortality of common eiders (Somateria mollissima) in the Dutch Wadden Sea, winter 1999/2000: starvation in a commercially exploited wetland of international importance. Biological Conservation 106(3): 303-317.

⁶⁰Camphuysen C.J., Heubeck M., Cox S., Bao R., Humple D., Abraham C. & Sandoval A. 2002. The Prestige oil spill in Spain. Atlantic Seabirds 4(3): 131-140.

⁶¹Camphuysen C.J. 2003. Seabirds and marine mammals off West Africa. Responses 2000 cruise report, Netherlands Institute for Sea Research, 6 January 2003, Texel.

⁶²Heubeck M., Camphuysen C.J., Bao Casal R., Humple D., Sandoval Rey A., Cadiou B. & Bräger S. 2003. Assessing the impact of major oil sills on seabird populations. Marine Pollution Bulletin in press.

⁶³Camphuysen C.J. 2002. Seabirds and marine mammals off South Georgia. CSR Consultancy Report 2002.01, CSR Consultancy, Oosterend, Texel, 46pp.

⁶⁴Camphuysen C.J., R. Bao, H. Nijkamp & M. Heubeck (eds) 2007. Handbook on Oil Impact Assessment. Report to DG Environment, European Commission, Grant Agreement 07.030900/2005/42907/SUB/A5, Version 1.0, Royal Netherlands Institute for Sea Research, Texel. Available online www.oiledwildlife.eu.

⁶⁵Camphuysen C.J., C. Smeenk, M.J. Addink, H. van Grouw & O.E. Jansen 2008. Cetaceans stranded in the Netherlands from 1998 to 2007. Lutra 51(2): 87-122.

⁶⁶Camphuysen C.J., H.J.P. Vercruijsse & A.L. Spaans 2011. Colony- and age-specific seasonal dispersal of Herring Gulls Larus argentatus breeding in The Netherlands. Journal of Ornithology 152: 849-868.

 ⁶⁷Shamoun-Baranes J., W. Bouten, C.J. Camphuysen & E. Baaij 2010. Riding the tide: intriguing observations of gulls resting at sea. Ibis 153: 411-415.

 ⁶⁸Shamoun-Baranes J., E. van Loon, R.S. Purves, B. Speckmann, D. Weiskopf & C.J. Camphuysen 2012. Analysis and visualization of animal movement. Biol. Lett. 8: 6-9.

 ⁶⁹Camphuysen C.J., J. Shamoun-Baranes, W. Bouten & S. Garthe 2012. Identifying ecologically important marine areas for seabirds using behavioural information in combination with distribution patterns. Biol. Conserv. 156: 22-29.

 ⁷⁰Bouten B., E.W. Baaij, J. Shamoun-Baranes & C.J. Camphuysen 2012. A flexible GPS tracking system for studying bird behaviour at multiple scales. J. Orn. DOI 10.1007

Publications 1977-2012

 Headers:

 Books

• Refereed contributions to books
• Special volumes and refereed reports
• Refereed papers
• Non-refereed papers
• Selected oral presentations
• Reports
• Posters

 All publications in reverse chronological order

 Books

Boyd I.J., Wanless S. & Camphuysen C.J. 2006 (eds). Top predators in Marine Ecosystems: monitoring change in upper trophic levels. Cambridge Univ. Press, Cambridge.

Camphuysen C.J. & Peet G. 2006. Walvissen en dolfijnen in de Noordzee. Fontaine Uitgevers, Kortenhoef.

Camphuysen C.J. & Peet G. 2006. Whales and dolphins of the North Sea. Fontaine Uitgevers, Kortenhoef.

Bijlsma R.G., Hustings F. & Camphuysen C.J. 2001. Schaarse en algemene vogels van Nederland. Avifauna van Nederland, 2. KNNV Uitgeverij Utrecht en GMB Uitgeverij, Haarlem, 497pp.

Refereed contributions to books

Zuur A.F. & C.J. Camphuysen 2012. Generalised additive models applied on northern gannets. In: Zuur A.F. (ed.) A beginner's guide to generalised additive models with R: 145-168. Highland Statistics, Newburgh.

Camphuysen C.J., B. Scott & S. Wanless 2006. Distribution and foraging interactions of seabirds and marine mammals in the North Sea: multi-species foraging assemblages and habitat-specific feeding strategies. In: Boyd I.J., Wanless S. & Camphuysen C.J. (eds) Top predators in Marine Ecosystems: monitoring change in upper trophic levels: 82-97. Cambridge Univ. Press, Cambridge.

Camphuysen C.J. 2001. Seabirds and fisheries interaction. In: Steele J.H., Thorpe S.A. & Turekian K.K. (eds) Encyclopedia of Ocean Sciences: 2677-2686. Academic Press, London.

Bourne W.R.P. & Camphuysen C.J. 2000. Seabirds. Chapter 114, In: Sheppard C. (ed.) Seas at the Millenium, an environmental evaluation: 105-116. Elsevier, Amsterdam.

Camphuysen C.J. & Garthe S. 2000. Seabirds and commercial fisheries: population trends of piscivorous seabirds explained? Chapter 11 In: Kaiser M.J. & Groot S.J. de (eds). Effects of fishing on non-target species and habitats: Biological, Conservation and Socio-Economic Issues: 163-184. Blackwell Science, Oxford.

Camphuysen C.J. 1997. Ivory Gull Pagophila eburnea. In: Hagemeijer J.M. & Blair M. (eds) The EBCC atlas of European breeding birds: their distribution and abundance: 348. T. & A.D. Poyser, London.

Camphuysen C.J. 1994. Ivory Gull Pagophila eburnea. In: Tucker G.M. & Heath M.F. (eds). Birds in Europe - their conservations status: 290-291. Birdlife Conservation Series No. 3, Birdlife International, Cambridge.

Camphuysen C.J. 1985. Zeetrektellingen. In: Hustings M.F.H., Kwak R.G.M., Opdam P.F.M. & Reijnen M.J.S.M. (eds). Vogelinventarisatie: 215-219. Pudoc, Wageningen.

Special volumes and refereed reports

Camphuysen C.J. & M.L. Siemensma 2011. Conservation plan for the Harbour Porpoise Phocoena phocoena in The Netherlands: towards a favourable conservation status. NIOZ Report 2011-07, Royal Netherlands Institute for Sea Research, Texel.

Haelters J. & C.J. Camphuysen 2009. The harbour porpoise (Phocoena phocoena L.) in the southern North Sea: Abundance, threats, research- and management proposals. Royal Belgian Institute of Natural Sciences (RBINS), department Management Unit of the North Sea Mathematical Models (MUMM) & Royal Netherlands Institute for Sea Research (NIOZ) Project financed by IFAW (International Fund for Animal Welfare) - IFAW Internationaler Tierschutz-Fonds GmbH, IFAW, Brussels.

Camphuysen C.J. 2007. Chronic oil pollution in Europe, a status report. Report Royal Netherlands Institute for Sea Research, commissioned by International Fund for Animal Welfare (IFAW), Brussels, 85pp.

Camphuysen C.J., R. Bao, H. Nijkamp & M. Heubeck (eds) 2007. Handbook on Oil Impact Assessment. Report to DG Environment, European Commission, Grant Agreement 07.030900/2005/42907/SUB/A5, Version 1.0, Royal Netherlands Institute for Sea Research, Texel. Available online www.oiledwildlife.eu.

Leopold M.F. & C.J. Camphuysen 2006. Bruinvisstrandingen in Nederland in 2006: Achtergronden, leeftijdsverdeling, sexratio, voedselkeuze en mogelijke oorzaken. IMARES Rapport C083/06, NIOZ Report 2006-5, Wageningen IMARES, Texel en Koninklijk Nederlands Instituut voor Onderzoek der Zee, Texel.

Camphuysen C.J. (ed.) 2005. Understanding marine foodweb processes: an ecosystem approach to sustainable sandeel fisheries in the North Sea. IMPRESS Final Report, EU Project #Q5RS-2000-30864, Royal Netherlands Institute for Sea Research, Texel.

Camphuysen C.J., A.D. Fox, M.F. Leopold & I.K. Petersen 2004. Towards standardised seabirds at sea census techniques in connection with environmental impact assessments for offshore wind farms in the U.K. Report commissioned by COWRIE for the Crown Estate, London. Royal Netherlands Institute for Sea Research, Texel, 38pp.

Jenni L. & C.J. Camphuysen (eds) 2001. Bird ringing 100 years. Proc. int. conf. Helgoland, 29 Sep - 3 Oct 1999. Ardea 89 (special issue): 1-252.

Camphuysen C.J., Calvo B., Durinck J., Ensor K., Follestad A., Furness R.W., Garthe S., Leaper G., Skov H., Tasker M.L. & Winter C.J.N. 1995. Consumption of discards by seabirds in the North Sea. Final report to the European Comm., study contr. BIOECO/93/10, NIOZ-Report 1995-5, Netherlands Institute for Sea Research, Texel, 202+LVIIIpp.

Stone C.J., Webb A., Barton C., Ratcliffe N., Reed T.C., Tasker M.L., Camphuysen C.J. & Pienkowski M.W. 1995. An atlas of seabird distribution in north-west European waters. Joint Nature Conservation Committee, Peterborough, 326pp.

Camphuysen C.J. & Leopold M.F. 1994. Atlas of seabirds in the southern North Sea. IBN Research report 94/6, NIOZ-Report 1994-8, Institute for Forestry and Nature Research, Netherlands Institute for Sea Research and Dutch Seabird Group, Texel.

Camphuysen C.J., Ensor K., Furness R.W., Garthe S., Hüppop O., Leaper G., Offringa H. & Tasker M.L. 1993. Seabirds feeding on discards in winter in the North Sea. Final report to the European Comm., study contr. 92/3505, NIOZ-Report 1993-8, Netherlands Institute for Sea Research, Texel.

Camphuysen C.J. & Franeker J.A. van 1992. The value of beached bird surveys in monitoring marine oil pollution. Techn. Rapport Vogelbescherming 10, Vogelbescherming Nederland, Zeist, 191pp.

Camphuysen C.J. 1991. Hvalguide - Veiledning i bestemmelse av hval i de Nordøst-Atlantiske havområder. Norsk Inst. Naturforsk., Trondheim NINA Temahefte 1: 1-41.

Camphuysen C.J. 1990. Fish stocks, fisheries and seabirds in the North Sea. Techn. Rapport Vogelbescherming nr. 5, Vogelbescherming Nederland, Zeist.

Camphuysen C.J. 1989. Beached Bird Surveys in the Netherlands 1915-1988; Seabird Mortality in the southern North Sea since the early days of Oil Pollution. Techn. Rapport Vogelbescherming 1, Werkgroep Noordzee, Amsterdam 322pp.

Camphuysen C.J. 1989. Olieslachtoffertellingen aan de Nederlandse kust, 1915-1988. Techn. Rapport Vogelbescherming 2, Werkgroep Noordzee, Amsterdam 21pp.

Camphuysen C.J. & Dijk J.van 1983. Zee- en kustvogels langs de Nederlandse kust, 1974-79. Limosa special issue 56(3): 81-230.

Refereed papers

Bouten B., E.W. Baaij, J. Shamoun-Baranes & C.J. Camphuysen 2012. A flexible GPS tracking system for studying bird behaviour at multiple scales. J. Orn. DOI 10.1007/s10336-012-0908-1.

Camphuysen C.J. & A. Gronert 2012. Apparent survival and fecundity of sympatric Lesser Black-backed Gulls and Herring Gulls with contrasting population trends. Ardea 100: 113-122.

Camphuysen C.J., J. Shamoun-Baranes, W. Bouten & S. Garthe 2012. Identifying ecologically important marine areas for seabirds using behavioural information in combination with distribution patterns. Biol. Conserv. 156: 22-29.

Embling C.B., J. Illian, E. Armstrong, J. van der Kooij, J. Sharples, C.J. Camphuysen & B.E. Scott 2012. Investigating fine scale spatio-temporal predator-prey patterns in dynamic marine ecosystems: a functional data analysis approach. J. Appl. Ecol. doi: 10.1111/j.1365-2664.2012.02114.x.

Shamoun-Baranes J., E. van Loon, R.S. Purves, B. Speckmann, D. Weiskopf & C.J. Camphuysen 2012. Analysis and visualization of animal movement. Biol. Lett. 8: 6-9.

Hallgrimsson G.T., H. Gunnarsson, O. Torfason, R-J. Buijs & C.J. Camphuysen 2012. Migration pattern of Icelandic Lesser Black-backed Gulls Larus fuscus graellsii: indications of a leap-frog system. Journal of Ornithology DOI 10.1007/s10336-012-0816-4.

Camphuysen C.J. 2011. Recent trends and spatial patterns in nearshore sightings of harbour porpoises (Phocoena phocoena) in the Netherlands (Southern Bight, North Sea), 1990-2010. Lutra 54(1): 37-44.

Camphuysen C.J. (Kees), J. Shamoun-Baranes, S. Garthe & W. Bouten The identification of offshore marine areas of ecological importance for seabirds based on recorded behaviour during ship-based transects or with data loggers. Biological Conservation submitted Feb 2011

Bartolomé C., Maside X., Camphuysen K. (C.J.), Heubeck M., Bao R. 2011. Multilocus population analysis of Common Loon (Gavia immer) mtDNA reveals low genetic diversity and lack of differentiation across the species breeding range. Organisms Diversity and Evolution 11: 307-316.

Garthe S., Camphuysen C.J., Fureness R.W. & Kubetzki U. [submitted, under review] Food consumption by seabirds in the North Sea. Ices Journal of Marine Science

Camphuysen C.J. 2011. Northern gannets Morus bassanus in the North Sea: foraging distribution and feeding techniques around the Bass Rock. Brit. Birds 104: 60-76.

Camphuysen C.J., H.J.P. Vercruijsse & A.L. Spaans 2011. Colony- and age-specific seasonal dispersal of Herring Gulls Larus argentatus breeding in The Netherlands. Journal of Ornithology DOI: 10.1007/s10336-011-0669-2.

Camphuysen C.J., S. Schouten & A. Gronert 2011. Mystery spill of Polyisobutylene (C4H8)n off the Dutch coast affecting seabirds in March 2010. Seabird 23: 143-145

Camphuysen, C.J., Bao, R., Fortin M., Roselaar, C.S. & Heubeck, M. 2011. Post-mortem examination of Great Northern Divers Gavia immer killed in the Prestige oil spill, Galicia, Spain 2002-3. Seabird 23: 53-65.

Camphuysen C.J. 2011. Seabirds and chronic oil pollution: self-cleaning properties of gulls, Laridae, as revealed from colour-ring sightings. Marine Pollution Bulletin 62: 514-519.

Wojczulanis-Jakubas K., D. Jakubas, J. Welcker, A.M.A. Harding, N.J. Karnovsky, D. Kidawa, H. Steen, L. Stempniewicz & C.J. Camphuysen 2010. Body size variation of a high-arctic seabird: the dovekie (Alle alle). Polar Biology 34: 847-854

Shamoun-Baranes J., W. Bouten, C.J. Camphuysen & E. Baaij 2010. Riding the tide: intriguing observations of gulls resting at sea. Ibis 153: 411-415.

Scott B.E., J. Sharples, O.N. Ross, J. Wang, G.J. Pierce & C.J. Camphuysen 2010. Sub-surface hotspots in shallow seas: fine scale limited locations of marine top-predator foraging habitat indicated by tidal mixing and sub-surface chlorophyll. Mar. Ecol. Progr. Ser. 408: 207-226.

Haelters J., F. Kerckhof & C.J. Camphuysen 2010. The first historic record of a humpback whale (Megaptera novaeangliae) from the Low Countries (Southern Bight of the North Sea). Lutra 53: 93-100.

Camphuysen C.J. & A. Gronert 2010. De broedbiologie van Zilver- en Kleine Mantelmeeuwen op Texel, 2006-2010. Limosa 83: 145-159.

Camphuysen C.J. 2010. Declines in oil-rates of stranded birds in the North Sea highlight spatial patterns in reductions of chronic oil pollution. Mar. Pollut. Bull. 60: 1299-1306.

Camphuysen C.J., P. de Boer, W. Bouten, A. Gronert & J. Shamoun-Baranes 2010. Mammalian prey in Laridae: increased predation pressure on mammal populations expected. Lutra 53(1): 5-20.

Camphuysen C.J. 2010. Intra-specific competition and other factors leading to far-ranging foraging flights in Northern Gannets Morus bassanus. British Birds [reviewed, provisionally accepted, awaits re-submission]

Camphuysen C.J., C. Smeenk, M.J. Addink, H. van Grouw & O.E. Jansen 2008. Cetaceans stranded in the Netherlands from 1998 to 2007. Lutra 51(2): 87-122.

Camphuysen C.J. & M.F. Leopold 2008. Drieteenmeeuw vestigt zich op meerdere platforms in Nederlandse wateren. Limosa 80: 153-156.

Barrett R.T., C.J. Camphuysen, T. Anker-Nilssen, J.W. Chardine, R.W. Furness, S. Garthe, O. Hüppop, M.F. Leopold, W.A. Montevecchi, R.R. Veit 2007. Diet studies of seabirds: a review and recommendations. ICES J. Mar. Sc. 64: 1675-1691.

Camphuysen C.J. 2007. Foraging humpback whale (Megaptera novaeangliae) in the Marsdiep area (Wadden Sea), May 2007 and a review of sightings and strandings in the southern North Sea, 2003-2007. Lutra 50(1): 31-42.

Camphuysen C.J. 2007. Where two oceans meet: offshore interactions of Great-winged Petrels Pterodroma macroptera and Leach's Storm petrels Oceanodroma leucorhoa off southern Africa. J. Ornithol. 148: 333-346.

Camphuysen C.J., D.C. Camphuijsen & T.M. van Spanje 2006. Het voedsel van de Kleine Mantelmeeuwen van het Wormer- en Jisperveld. Limosa 78: 145-154.

Ens B.J., Kats R. & Camphuysen C.J. 2006. Waarom zijn Eiders niet massaal gestorven in de winter van 2005/2006? Limosa 79: 95-106.

Meij S.E.T. van der & C.J. Camphuysen 2006. Distribution and diversity of whales and dolphins (Cetacea) in the Southern North Sea: 1970-2005. Lutra 49(1): 3-28.

Camphuysen C.J. 2005. Assessing age and breeding origin of wrecked Little Auks Alle alle: the use of biometrics and a variable underwing pattern. Atlantic Seabirds 7(2): 49-70.

Camphuysen C.J. 2005. Seabirds at sea in summer in the NW North Sea. British Birds 98: 2-19.

Camphuysen C.J. 2005. The Tricolor oil spill: an incident that should have been prevented. Atlantic Seabirds 6(3): 81-84.

Camphuysen C.J. & F. de Vreeze 2005. De Drieteenmeeuw Rissa tridactyla als broedvogel in Nederland. Limosa 78(2): 65-74.

Camphuysen C.J. & J. van der Meer 2005. Wintering seabirds in Western Africa: foraging hot-spots off Western Sahara and Mauritania driven by upwelling and fisheries. African J. Mar. Sc. 27(2): 427-437.

Camphuysen C.J. & M.F. Leopold 2005. The Tricolor oil spill: characteristics of seabirds found oiled in The Netherlands. Atlantic Seabirds 6(3): 109-128.

Ouwehand J., Leopold M.F. & Camphuysen C.J. 2005. A comparative study of the diet of Guillemots Uria aalge and Razorbills Alca torda killed during the Tricolor oil incident in the south-eastern North Sea in January 2003. Atlantic Seabirds 6(3): 147-164.

Camphuysen C.J. 2004. Notes on seabirds 78. Deposition rates of carcasses on the beach in The Netherlands. Atlantic Seabirds 6(2): 79-80.

Camphuysen C.J. 2004. The return of the harbour porpoise (Phocoena phocoena) in Dutch coastal waters. Lutra 47(2): 113-122.

Camphuysen C.J. & Garthe S. 2004. Recording foraging seabirds at sea: standardised recording and coding of foraging behaviour and multi-species foraging associations. Atlantic Seabirds 6(1): 1-32.

Leopold M.F., Bruin C.J.W., Camphuysen C.J., Winter C.J.N. & Koks B.J. 2004. Waarom is de Visarend Pandion haliaetus in Nederland geen zeearend? Limosa 76: 129-140.

Camphuysen C.J. 2003. Characteristics of Atlantic Puffins Fratercula arctica wrecked in The Netherlands, January-February 2003. Atlantic Seabirds 5(1): 21-30.

Heubeck M., Camphuysen C.J., Bao R., Humple D., Sandoval A., Cadiou B., Bräger S. & Thomas T. 2003. Assessing the impact of major oil spills on seabird populations. Mar. Poll. Bull. 46: 900-902.

Camphuysen C.J. 2002. Post-fledging dispersal of Common Guillemots Uria aalge guarding chicks in the North Sea: the effect of predator presence and prey availability at sea. Ardea 90(1): 103-119.

Camphuysen C.J., C.M. Berrevoets, H.J.W.M. Cremers, A. Dekinga, R. Dekker, B.J. Ens, T.M. van der Have, R.K.H. Kats, T. Kuiken, M.F. Leopold, J. van der Meer & T. Piersma 2002. Mass mortality of common eiders (Somateria mollissima) in the Dutch Wadden Sea, winter 1999/2000: starvation in a commercially exploited wetland of international importance. Biological Conservation 106(3): 303-317.

Camphuysen C.J., Heubeck M., Cox S., Bao R., Humple D., Abraham C. & Sandoval A. 2002. The Prestige oil spill in Spain. Atlantic Seabirds 4(3): 131-140.

Desholm M., Christensen T.K., Scheiffarth G., Hario M., Andersson Å., Ens B., Camphuysen C.J., Nilsson L., Waltho C.M., Lorentsen S-H., Kuresoo A., Kats R.K.H., Fleet D.M. & Fox A.D. 2002. Status of the Baltic/Wadden Sea population of the Common Eider Somateria m. mollissima. Wildfowl 53: 167-203.

Underhill L.G., Crawford R.J.M. & Camphuysen C.J. 2002. Leach's storm petrels Oceanodroma leucorhoa off southern Africa: breeding and migratory status, and measurements and mass of the breeding population. Trans. Roy. Soc. S. Afr. 57: 43-46.

Camphuysen C.J. 2001. The distribution of Spectacled Petrels Procellaria conspicillata in the south-eastern Atlantic. Atlantic Seabirds 3(3): 113-124.

Camphuysen C.J. 2001. Northern Gannets Morus bassanus found dead in The Netherlands, 1970-2000. Atlantic Seabirds 3(1): 15-30.

Camphuysen C.J. & Heubeck M. 2001. Marine oil pollution and beached bird surveys: the development of a sensitive monitoring instrument. Environmental Pollution 112: 443-461.

Camphuysen C.J. & Van der Meer J. 2001. Pelagic distribution, moult and (sub-) specific status of Cory's Shearwaters Calonectris [d.] diomedea/borealis wintering off southern Africa. Marine Ornithology 29: 85-93.

Leopold M.F., Leeuwen P.W. van, Horn H. & Camphuysen C.J. 2001. Grote IJsduiker Gavia immer eet kleine visjes: een verdronken vogel geeft geheimen prijs. Limosa 73: 135-143.

Camphuysen C.J. 2000. Mass mortality of Common Eiders Somateria mollissima in the Wadden Sea, winter 1999/2000: food related parasite outbreak? Atlantic Seabirds 2(1): 47-48.

Camphuysen C.J. 2000. Seabirds drowned in fishing nets off Jan Mayen (Greenland Sea). Atlantic Seabirds 2(2): 87-91.

Camphuysen C.J. 2000. Zomerwaarnemingen van Noordse Pijlstormvogels Puffinus puffinus in Nederland. Limosa 73: 7-16.

Camphuysen C.J. & J. van der Meer 2000. Notes on the distribution of the Spectacled Petrel Procellaria conspicillata in the South Atlantic. Atlantic Seabirds 2(1): 13-18.

Tasker M.L., Camphuysen C.J., Cooper J., Garthe S., Montevecchi W.A. & Blaber S.J.M. 2000. The impacts of fishing on marine birds. ICES J. Mar. Sc. 57: 531-547.

Camphuysen C.J. 1999. Diurnal activity patterns and nocturnal group formation of wintering Common Murres in the central North Sea. Colonial Waterbirds 21(3): 406-413.

Camphuysen C.J. 1999. New feeding technique of Great Cormorants Phalacrocorax carbo sinensis at beam trawlers. Atlantic Seabirds 1(2): 85-90.

Camphuysen C.J. & A. Webb 1999. Multi-species feeding associations in North Sea seabirds: jointly exploiting a patchy environment. Ardea 87(2): 177-198.

Camphuysen C.J., Barreveld H., Dahlmann G. & Franeker J.A. van 1999. Seabirds in the North Sea demobilised and killed by polyisobutylene (C₄H₈)_n. Marine Pollution Bulletin 38(12): 1171-1176.

Camphuysen C.J. & Reid J.B. 1999. Trends in seabird systematics: recent sometimes conflicting decisions of BOURC and CSNA. Atlantic Seabirds 1(2): 92-94.

Phillips R.A., M.K. Petersen, K. Lilliendahl, J. Solmundsson, K.C. Hamer, C.J. Camphuysen & B. Zonfrillo 1999. The diet of the Northern Fulmar Fulmarus glacialis: reliance on commercial fisheries? Marine Biology 135: 159-170.

Camphuysen C.J. 1998. Beached bird surveys indicate decline in chronic oil pollution in the North Sea. Mar. Poll. Bull. 36(7): 519-526.

Camphuysen C.J. 1998. Het voorkomen van de Alk Alca torda in Nederlandse wateren. Limosa 71: 69-77.

Franeker J.A. van, Camphuysen C.J. & Mehlum F. 1998. The birds of Jan Mayen. Circumpolar Journal 13(2): 28-43.

Camphuysen C.J. & Garthe S. 1997. An evaluation of the distribution and scavenging habits of Northern Fulmars (Fulmarus glacialis) in the North Sea. ICES J. Mar. Sc. 54: 654-683.

Furness R.W. & Camphuysen C.J. 1997. Seabirds as monitors of the marine environment. ICES J. Mar. Sc. 54: 726-737.

Northridge S., Tasker M.L., Webb A., Camphuysen C.J. & Leopold M.F. 1997. White-beaked Lagenorhynchus albirostris and Atlantic white-sided L. acutus distributions in north-west European and U.S. North Atlantic waters. SC/48/SM44, Rep. Int. whal. Commn. 47: 797-805.

Camphuysen C.J., Ens B.J., Heg D., Hulscher J., Meer J. van der & Smit C.J. 1996. Oystercatcher winter mortality in The Netherlands: the effect of severe weather and food supply. Ardea 84a: 469-492.

Camphuysen C.J. & Winter C.J.N. 1996. Arctic Terns Sterna paradisaea in the central northern North Sea in July: offshore staging area for failed breeders? Seabird 18: 20-25.

Garthe S., Camphuysen C.J. & Furness R.W. 1996. Amounts of discards in commercial fisheries and their significance as food for seabirds in the North Sea. Mar. Ecol Progr. Ser. 136: 1-11.

Meer J. van der & Camphuysen C.J. 1996. Effect of observer differences on abundance estimates of seabirds from ship-based strip transect surveys. Ibis 138: 433-437.

Reijnders P.J.H., Leopold M.F., Camphuysen C.J., Heessen H.J.L. & Kastelein R.A. 1996. The status of harbour porpoises in Dutch waters and state of related research in The Netherlands: an overview. SC/47/SM41, Rep. Int. Whal. Commn. 46: 607-611.

Camphuysen C.J. 1995. Grauwe Pijlstormvogel Puffinus griseus en Noordse Pijlstormvogel P. puffinus in de zuidelijke Noordzee: een offshore perspectief. Limosa 68: 1-9.

Camphuysen C.J. 1995. Herring Gull Larus argentatus and Lesser Black-backed Gulls Larus fuscus feeding at fishing vessels in the breeding season: competitive scavenging versus efficient flying. Ardea 83(2): 365-380.

Camphuysen C.J., Heessen H.J.L. & Winter C.J.N. 1995. Distant feeding and associations with cetaceans of Gannets Morus bassanus from Bass Rock, May 1994. Seabird 17: 36-43.

Camphuysen C.J. & Winter C.J.N. 1995. Feeding Fin Whales Balaenoptera physalus in the North Sea. Lutra 38: 81-84.

Camphuysen C.J. 1994. Flatfish selection by Herring Gulls Larus argentatus and Lesser Black-backed Gulls Larus fuscus scavenging at commercial beamtrawlers in the southern North Sea. Neth. J. Sea Res. 32(1): 91-98.

Camphuysen C.J. 1994. The Harbour Porpoise Phocoena phocoena in the southern North Sea, II: a come-back in Dutch coastal waters? Lutra 37(1): 54-61.

Dahlmann G., Timm D., Averbeck C., Camphuysen C.J. & Skov H. 1994. Oiled seabirds - Comparative investigations on oiled seabirds and oiled beaches in the Netherlands, Denmark and Germany (1990-1993). Mar. Poll. Bull. 28: 305-310.

Camphuysen C.J. & Leopold M.F. 1993. The Harbour Porpoise Phocoena phocoena in the southern North Sea, particularly the Dutch sector. Lutra 36(1): 1-24.

Camphuysen C.J. 1992. Zeevogelstrandingen op de Nederlandse kust: 26 jaar een vinger aan de pols (1965-1991). Limosa 66(1): 1-16.

Camphuysen C.J. 1990. Biometrics of auks at Jan Mayen. Seabird 12: 7-10.

Camphuysen C.J. 1989. Diurnal rhythm of the Fulmar Fulmarus glacialis in the arctic summer. Dansk Orn. Foren. Tidsskr. 83: 85-86.

Camphuysen C.J. 1989. Vondst van een noordelijke Zeekoet Uria aalge hyperborea in Nederland. Limosa 62(1): 47-48.

Camphuysen C.J. & Derks P.J.T. 1989. Voorkomen en sterfte van de Fuut Podiceps cristatus voor de Nederlandse kust, 1974-86. Limosa 62: 57-62.

Camphuysen C.J. & IJzendoorn E.J. van 1988. Influx of Pomarine Skua in northwestern Europe in autumn 1985. Dutch Birding 10(2): 66-70.

Camphuysen C.J. & IJzendoorn E.J. van 1988. Invasie van Middelste Jagers in Nederland in november 1985. Dutch Birding 10(2): 54-65.

Camphuysen C.J. 1986. Vondsten van Kleine Alken Alle alle en Papegaaiduikers Fratercula arctica langs de Nederlandse kust. Limosa 59(3): 138-141.

Franeker J.A. van, Camphuysen C.J. & Mehlum F. 1986. Status over Jan Mayens fugler. Vår Fuglefauna 9(3): 145-158.

Camphuysen C.J. 1982. Vondst van een Kuhls Pijlstormvogel Calonectris diomedea. Limosa 55(3): 99-100.

Camphuysen C.J. & Maas F.J. 1982. Zeevogels in Nederland in 1978. Limosa 55: 17-22.

Non-refereed papers

Camphuysen C.J., R. Dekker & A. Gronert 2012. Zilvermeeuwen op strandhoofden langs duin en dijk: een verguisde vogelsoort in een bedreigd habitat. Tussen Duin en Dijk 10(1): 4-7.

Camphuysen C.J. & M. Heubeck 2012. Post-mortale migratie van een Noordse Stormvogel over de Noordzee. Sula 25(1): 45-46.

Camphuysen C.J. 2010. Olieslachtoffers op het strand: wordt de zee nu werkelijk schoner?. Tussen Duin & Dijk 9(4): 29-31.

Camphuysen C.J. 2010. Walvissen in Noord-Holland, vroeger en nu. Tussen Duin en Dijk 9(4): 14-18.

Kleinwee M. van & C.J. Camphuysen 2010. R